Peristalsis is Regulated by the CNS and Enteric Nervous System for Food Movement
- The movement of food through the digestive system is regulated by both voluntary and involuntary control mechanisms.
- Imagine taking a bite of your favorite food.
- As you swallow, the food travels down your esophagus, through your stomach, and into your intestines.
- This journey is powered by peristalsis, a series of muscle contractions that move food through the digestive tract.
Peristalsis
Peristalsis is a wave-like contraction of smooth muscles that pushes food through the digestive tract.
The Role of the Central Nervous System (CNS)
The CNS, comprising the brain and spinal cord, initiates and controls voluntary actions in the digestive process, such as swallowing and egestion.
1. Swallowing: A Two-Phase Process
- Voluntary Phase: You consciously push food to the back of your mouth using your tongue.
- Involuntary Phase: Once food reaches the pharynx, sensory receptors trigger a reflex controlled by the brainstem, moving food into the esophagus.
Swallowing is a rare example of an action that begins voluntarily but quickly becomes involuntary.
2. Egestion: Voluntary Control of Defecation
- Rectal Filling: As the rectum fills with feces, stretch receptors send signals to the CNS.
- Voluntary Decision: The brain decides whether to relax the external anal sphincter, allowing defecation.
Infants lack voluntary control over defecation, which is why toilet training is necessary.
The Enteric Nervous System (ENS):
Enteric nervous system
The enteric nervous system, often called the “second brain,” responsible for controlling involuntary muscle contractions in the digestive tract.
- The ENS is a complex network of neurons embedded in the walls of the digestive tract.
- It operates independently of the CNS, controlling involuntary processes like peristalsis.
How the ENS Controls Peristalsis
- Muscle Layers: The gut wall has two layers of smooth muscle:
- Circular Muscle: Contracts to narrow the gut.
- Longitudinal Muscle: Contracts to shorten the gut.
- Coordinated Contractions: The ENS coordinates these muscles to create wave-like contractions that push food forward.
- In the esophagus, peristalsis moves food quickly to the stomach.
- In the intestines, it progresses more slowly, allowing time for digestion and absorption.
Autonomy of the ENS
- The ENS can function without CNS input, thanks to its intrinsic microcircuits.
- However, it remains connected to the CNS, which can modulate its activity.
- Don’t confuse the ENS with the CNS.
- The ENS operates independently but can be influenced by the CNS.
Integration of the CNS and ENS
While the ENS handles most digestive activities, the CNS plays a crucial role in initiating and terminating the process.
1. Initiation: Swallowing
- Begins voluntarily under CNS control.
- Transitions to ENS control for peristalsis in the esophagus.
2. Termination: Egestion
- The ENS manages rectal contractions.
- The CNS controls the external anal sphincter, allowing voluntary defecation.
Think of the CNS as the manager that starts and ends the process, while the ENS is the dedicated worker handling the tasks in between.
Theory of Knowledge- How does the independence of the ENS challenge our understanding of the nervous system?
- Could it be considered a "second brain"?
- Which parts of the digestive system are controlled by the CNS and which are controlled by the ENS?
- Describe the process of peristalsis and the role of the ENS in this process



