Species diversity
Species diversity is the variety of species in an ecosystem, including both species richness (the number of species) and species evenness (the relative abundance of each species).
- Species diversity is a key measure of biodiversity in an ecosystem, influenced by species richness and species evenness.
- These two factors together determine how balanced and functionally stable a community is.
Components of Species Diversity
Species Richness
Species richness
Species richness is the total number of different species present in a community.
- It provides a simple count of how many different types of organisms are present.
- Although it gives an overview of biodiversity, it does not reveal how evenly individuals are distributed across species.
- Higher species richness means greater variety in an ecosystem, leading to more ecological roles and interactions.
A coral reef with 100 species of fish has higher species richness than a lake with only 10 fish species.
Species Evenness
Species evenness
Species evenness is the relative abundance of each species in a community.
- Describes how uniformly individuals are distributed among the different species in a community.
- High evenness means that species have similar population sizes, suggesting a stable and balanced ecosystem.
- Low evenness occurs when one or a few species dominate, indicating lower stability and resilience.
A forest where all tree species have similar population sizes has high evenness, while a forest dominated by only one tree species has low evenness.
- Species richness and evenness together determine the overall species diversity in an ecosystem.
- While richness increases variety, evenness ensures balance, both of which contribute to ecosystem stability, resilience, and long-term biodiversity conservation.
Significance of Richness and Evenness for Biodiversity
- High Richness, Low Evenness (Unstable Ecosystem)
- A large number of species exist, but a few species dominate.
- Low evenness can lead to imbalances where dominant species outcompete others, reducing overall biodiversity.
- Low Richness, High Evenness (Limited Diversity, More Stability)
- Fewer species present, but they exist in similar proportions, meaning no single species outcompetes the rest.
- More stable than a low-evenness system, but still vulnerable to disturbances due to low species variety.
- High Richness, High Evenness (Most Stable and Resilient Ecosystem)
- Diverse species and balanced populations create a strong, resilient ecosystem.
- Supports high productivity, efficient resource use, and greater resistance to environmental changes.
Importance for Ecosystem Stability and Functioning
- Higher richness provides functional redundancy: multiple species can perform the same ecological role, preventing collapse if one species declines.
- Higher evenness prevents dominance: ensures no single species takes over, keeping the ecosystem balanced.
- Diverse and even communities are more resilient: ecosystems with both high richness and evenness recover faster from disturbances like climate change, fires, or disease.
- How does the way we measure species diversity influence conservation decisions?
- Are there ethical implications in prioritizing certain ecosystems or species over others?
Simpson’s Reciprocal Index
Simpson's reciprocal index
Simpson’s Reciprocal Index (D) is a quantitative measure of species diversity, used to: compare biodiversity between different ecosystems, monitor changes in biodiversity over time within a specific area and assess ecosystem health,
- Biodiversity can be quantified using mathematical indices that combine richness and evenness.
- The most widely used is Simpson’s Reciprocal Index (D), which provides a single value to compare ecosystems or track changes over time.
- Simpson’s reciprocal index provides:
- A quantitative measure of biodiversity.
- A way to compare ecosystems with similar species types.
- A method to monitor biodiversity changes in a single ecosystem over time.
The formula for Simpson’s Reciprocal Index
$$D = \frac{N(N - 1)}{\sum n(n - 1)}$$
Where:
- D is Diversity index (higher values = greater diversity).
- N is Total number of individuals in the sample (sum of all species counted).
- n is Number of individuals of a single species.
- Σ is Summation symbol (sum of all species calculations).
You are not required to memorize the formula, but you should understand what the symbols represent:
- Higher D value: greater diversity (many species, evenly distributed).
- Lower D value (close to 1): low diversity (dominated by few species).
- D = 1: only one species present.
Sampling Procedures for Calculating Simpson’s Index
- Choose an appropriate sampling area
- Select two or more sites for comparison (e.g., forest vs. grassland).
- Ensure sites are similar in size and environmental conditions.
- Use a standardized sampling method
- Quadrats (e.g., 1m² for plants, 10m² for trees).
- Transects (line or belt transects for zonation studies).
- Pitfall traps or sweep nets (for insects and small animals).
- Record species abundance
- Count the number of individuals per species.
- Repeat for multiple quadrats to improve accuracy.
Example Calculation of Simpson’s Reciprocal Index
Collected Data from Two Sample Sites (Same Ecosystem, Different Areas)
| Species | Site A (n) | Site B (n) |
|---|---|---|
| Oak tree | 12 | 20 |
| Pine tree | 8 | 25 |
| Birch tree | 10 | 5 |
| Maple tree | 6 | 10 |
| Total (N) | 26 | 60 |
Step 1: Calculate $N(N−1)$
- Site A: 36(36−1)=1260
- Site B: 60(60−1)=3540
Step 2: Calculate $∑n(n−1)$
- For Site A: (12(11)+8(7)+10(9)+6(5)) = (132+56+90+30)= 308
- For Site B: (20(19)+25(24)+5(4)+10(9)) = (380+600+20+90)=1090
Step 3: Calculate $D$
- For Site A: D = 1260/308 = 4.09
- For Site B: D = 3540/1090 = 3.25
Interpretation
- Site A has higher diversity ($D = 4.09$), meaning species are more evenly distributed.
- Site B has lower diversity ($D = 3.25$), suggesting one species may be more dominant.
Global and Regional Biodiversity Knowledge
- Conserving biodiversity requires comprehensive data collection at global, regional, and local levels.
- This knowledge guides management strategies and helps identify conservation priorities.
Global Data Collection
- International agencies such as the IUCN and WWF maintain global biodiversity databases.
- The IUCN Red List classifies species by extinction risk (e.g., Least Concern, Endangered, Critically Endangered).
- The WWF identifies and monitors biodiversity hotspots worldwide.
- These databases help governments and NGOs coordinate conservation actions and set priorities.
The IUCN lists the Kinabalu birdwing butterfly as vulnerable due to habitat loss on Mount Kinabalu, Malaysia.
Regional and Local Data Collection
- Government agencies (e.g., Natural England, Forest Survey of India).
- NGOs and voluntary organizations (e.g., Wildlife Trusts, BirdLife International).
- Citizen science projects, where volunteers record species observations (e.g., eBird, Big Butterfly Count).
- Indigenous and local communities, trained as parabiologists, contribute traditional ecological knowledge for monitoring and conservation.
Indigenous rangers in Australia monitor populations of threatened marsupials using traditional tracking skills combined with scientific methods.
The Kinabalu Birdwing Butterfly, Borneo
- Mount Kinabalu’s montane cloud forests are home to the Kinabalu birdwing butterfly, an endemic species dependent on the Aristolochia vine.
- Habitat loss and illegal collection have threatened the species.
- Conservation initiatives involve citizen science monitoring, indigenous ranger programs, and habitat protection.
- The butterfly’s data help scientists understand species distribution, seasonal activity, and reproductive patterns, supporting habitat-specific management.
Conservation Applications of Biodiversity Data
- Protected Areas: Identify biodiversity hotspots for national parks or reserves.
- Restoration: Track changes after reforestation or pollution mitigation.
- Policy Decisions: Support national biodiversity strategies aligned with the UN Sustainable Development Goals (SDG 14 & 15).
- Sustainable Use: Inform community-based resource management (fisheries, forestry).
- How does the integration of indigenous knowledge challenge traditional scientific approaches?
- What are the strengths and limitations of each?
- Define species richness and species evenness and explain how they contribute to overall diversity.
- Describe how Simpson’s reciprocal index is calculated and interpreted.
- Why can’t diversity indices be compared across entirely different ecosystems?
- How does biodiversity data support effective conservation and sustainable management at global and regional scales?


