Species diversity
Species diversity is the variety of species in an ecosystem, including both species richness (the number of species) and species evenness (the relative abundance of each species).
- Species diversity is a key measure of biodiversity in an ecosystem, influenced by species richness and species evenness.
- These two factors together determine how balanced and functionally stable a community is.
Components of Species Diversity
Species Richness
Species richness
Species richness is the total number of different species present in a community.
- It provides a simple count of how many different types of organisms are present.
- Although it gives an overview of biodiversity, it does not reveal how evenly individuals are distributed across species.
- Higher species richness means greater variety in an ecosystem, leading to more ecological roles and interactions.
A coral reef with 100 species of fish has higher species richness than a lake with only 10 fish species.
Species Evenness
Species evenness
Species evenness is the relative abundance of each species in a community.
- Describes how uniformly individuals are distributed among the different species in a community.
- High evenness means that species have similar population sizes, suggesting a stable and balanced ecosystem.
- Low evenness occurs when one or a few species dominate, indicating lower stability and resilience.
A forest where all tree species have similar population sizes has high evenness, while a forest dominated by only one tree species has low evenness.
Note- Species richness and evenness together determine the overall species diversity in an ecosystem.
- While richness increases variety, evenness ensures balance, both of which contribute to ecosystem stability, resilience, and long-term biodiversity conservation.
Significance of Richness and Evenness for Biodiversity
- High Richness, Low Evenness (Unstable Ecosystem)
- A large number of species exist, but a few species dominate.
- Low evenness can lead to imbalances where dominant species outcompete others, reducing overall biodiversity.
- Low Richness, High Evenness (Limited Diversity, More Stability)
- Fewer species present, but they exist in similar proportions, meaning no single species outcompetes the rest.
- More stable than a low-evenness system, but still vulnerable to disturbances due to low species variety.
- High Richness, High Evenness (Most Stable and Resilient Ecosystem)
- Diverse species and balanced populations create a strong, resilient ecosystem.
- Supports high productivity, efficient resource use, and greater resistance to environmental changes.
Importance for Ecosystem Stability and Functioning
- Higher richness provides functional redundancy: multiple species can perform the same ecological role, preventing collapse if one species declines.
- Higher evenness prevents dominance: ensures no single species takes over, keeping the ecosystem balanced.
- Diverse and even communities are more resilient: ecosystems with both high richness and evenness recover faster from disturbances like climate change, fires, or disease.
- How does the way we measure species diversity influence conservation decisions?
- Are there ethical implications in prioritizing certain ecosystems or species over others?
Simpson’s Reciprocal Index
Simpson's reciprocal index
Simpson’s Reciprocal Index (D) is a quantitative measure of species diversity, used to: compare biodiversity between different ecosystems, monitor changes in biodiversity over time within a specific area and assess ecosystem health,
- Biodiversity can be quantified using mathematical indices that combine richness and evenness.
- The most widely used is Simpson’s Reciprocal Index (D), which provides a single value to compare ecosystems or track changes over time.
- Simpson’s reciprocal index provides:
- A quantitative measure of biodiversity.
- A way to compare ecosystems with similar species types.
- A method to monitor biodiversity changes in a single ecosystem over time.
The formula for Simpson’s Reciprocal Index
$$D = \frac{N(N - 1)}{\sum n(n - 1)}$$
Where:
- D is Diversity index (higher values = greater diversity).
- N is Total number of individuals in the sample (sum of all species counted).
- n is Number of individuals of a single species.
- Σ is Summation symbol (sum of all species calculations).
You are not required to memorize the formula, but you should understand what the symbols represent:
Note- Higher D value: greater diversity (many species, evenly distributed).
- Lower D value (close to 1): low diversity (dominated by few species).
- D = 1: only one species present.


