Zonation
Zonation refers to the gradual change in species composition along an environmental gradient.
- Zonation occurs as abiotic and biotic conditions vary across space, resulting in distinct zones where particular species dominate.
- It represents a spatial pattern (not temporal like succession) and is often observed in ecosystems such as rocky shores, sand dunes, mountains, and forests.
- It describes how biotic communities (plants, animals, and microorganisms) change in response to abiotic factors such as:
- Elevation (altitude)
- Latitude
- Tidal level
- Soil horizons
- Distance from a water source
- Temperature, moisture, and light intensity
- As you ascend a mountain for hiking, the vegetation may change from tropical forest to temperate woodland to alpine meadow.
- This gradual shift in community composition illustrates zonation.
Factors Influencing Zonation
Changes in Abiotic Factors
- Elevation (altitude): Temperature and oxygen levels decrease, changing vegetation types from tropical forests at low altitude to alpine tundra higher up.
- Latitude: Sunlight intensity and temperature vary from the equator to the poles, influencing global biome zonation.
- Tidal level: In coastal ecosystems, exposure to air, salinity, and wave action varies between high and low tide zones.
- Soil composition: Soil pH, salinity, and nutrient availability differ between areas, creating patterns in plant distribution.
- Distance from water source: Moisture availability affects vegetation density and type, as lush vegetation near rivers transitions to arid desert farther away.
On a mountain slope, zonation can be observed as tropical forests give way to montane forests, grasslands, and eventually alpine tundra as altitude increases.
Changes in Biotic Factors
- Competition: Species compete for light, space, or nutrients, influencing their zonal distribution.
- Predation: Predators may restrict the range of prey species to specific zones.
- Mutualism: Certain plants may only grow in areas where their symbiotic species (e.g., pollinators or mycorrhizae) exist.
- Always connect zonation patterns to both abiotic and biotic interactions.
- For example, seaweed zonation depends on sunlight exposure (abiotic) and grazing snails (biotic).
Examples of Zonation
1. Rocky Shore Zonation
- The rocky intertidal zone provides a clear, observable example of zonation.
- Conditions vary sharply from the upper shore (exposed, dry, salty) to the lower shore (submerged, stable, oxygen-rich).
- Species distribution:
- Spray zone: Exposed to air most of the time; inhabited by lichens and periwinkles.
- High tide zone: Alternates between wet and dry; barnacles and limpets dominate.
- Middle tide zone: Submerged for longer periods; mussels, seaweed, and chitons thrive.
- Low tide zone: Constantly submerged; anemones, sea urchins, and kelp are abundant.
On rocky shores in the UK, channel wrack dominates the upper shore due to its resistance to desiccation, while kelp thrives in the lower shore where constant submersion prevents drying.
2. Mountain Ecosystems
- As altitude increases:
- Temperature decreases and oxygen levels drop.
- Plant communities transition from tropical forest → montane forest → alpine grasslands → lichens and mosses.
- Each band supports species adapted to that altitude’s conditions.
On Mount Kilimanjaro, zonation occurs from dense rainforests at the base to alpine tundra near the summit, a clear example of how temperature and moisture gradients determine vegetation zones.
3. Coastal Dune Zonation
- Sand dune systems exhibit gradual changes from the shore inland:
- Embryo dunes: Closest to the shore; few species (e.g., sand couch, sea rocket).
- Foredunes: Dominated by marram grass that stabilizes the sand.
- Fixed dunes: Contain shrubs and herbs like sea buckthorn and wild rose.
- Dune slacks: Low-lying areas between dunes; moisture supports diverse flora.
- Wooded dunes: Farther inland, trees like oak and birch dominate.
North Sea Coast of the Netherlands
- Gradual zonation from bare sand near the coast to forest inland.
- Factors changing along the gradient: decreasing salinity, wind speed, and sand movement.
- Vegetation progression:
- Pioneer zone: sand couch and sea rocket.
- Marram grass zone: stabilizes dunes.
- Shrub zone: sea buckthorn, wild rose.
- Woodland zone: birch, oak, and sycamore.
Scales of Zonation
- Local Scale:
- Observed within ecosystems (e.g., rocky shore, sand dunes).
- Determined by microclimate and soil differences.
- Global Scale:
- Seen across continents (e.g., from tropical to polar biomes).
- Determined by latitude, temperature, and precipitation.
Zonation can occur horizontally (across a beach) or vertically (up a mountain slope).
Measurements and Investigation of Zonation
To study biotic (living organisms) and abiotic (non-living) factors that influence this distribution, transects and kite diagrams can be used to collect and visualize data.
Using Transects to Study Zonation
Transects
A transect is a straight line or path across an environmental gradient along which samples are taken at regular intervals.
- A transect is a line or path laid out across an environmental gradient along which ecological data (biotic and abiotic) are collected at regular intervals.
- It allows researchers to quantify how species distribution changes with variations in environmental conditions.
Types of Transects
- Line Transect:
- A straight line marked along a gradient (e.g., from high to low tide).
- Organisms touching the line are recorded.
- Belt Transect:
- A strip (e.g., 1 m wide) along the line where all species within the belt are recorded.
- Provides more quantitative data on abundance.
- Interrupted Transect:
- Sampling occurs at fixed intervals (e.g., every 1 m).
- Useful when continuous sampling is impractical.
A belt transect from the seashore inland can reveal how salt-tolerant plants decrease as salinity drops and soil moisture stabilizes.
Investigating Zonation Using a Transect
Materials
- Measuring tape (for transect)
- Quadrat frame
- Soil thermometer, pH meter, and light meter
- Field notebook or digital data sheet
- Graph paper or spreadsheet (for kite diagrams)
Procedure
- Select a suitable environmental gradient (e.g., a rocky shore, sand dune, or forest edge).
- Lay a transect line perpendicular to the gradient (e.g., from low tide to high tide).
- Mark intervals (e.g., every 1 or 2 meters).
- At each interval:
- Measure abiotic variables (temperature, pH, salinity, etc.).
- Place a quadrat and record all species and their abundance.
- Record data systematically in a table for analysis.
- Plot results using kite diagrams to visualize zonation.
Measuring Abiotic and Biotic Factors
| Abiotic Factors | Measurement | Influence on Zonation |
|---|---|---|
| Temperature | Thermometer | Affects enzyme activity and species tolerance |
| Light intensity | Light meter | Determines photosynthesis rate |
| Soil pH | pH meter | Influences plant nutrient uptake |
| Salinity | Conductivity meter | Limits species based on salt tolerance |
| Moisture | Soil moisture probe | Affects plant water availability |
| Elevation | Altimeter | Changes temperature and pressure with height |
Biotic factors measured
- Species richness (number of species present)
- Species abundance (number of individuals per species)
- Percentage cover (using quadrats)
- Dominant species or communities
Kite Diagrams: Visualizing Zonation Data
Kite diagram
A kite diagram is a graphical representation of species distribution along a transect.
- Kite diagrams are used to display species distribution and abundance along a transect.
- The x-axis represents the distance along the transect, and the y-axis shows abundance (e.g., % cover).
- The width of the kite represents abundance, while its position along the transect shows where the species occurs.
- It shows:
- Species abundance at different points.
- Environmental changes along a gradient.
- Symmetrical "kite-like" shapes for each species.
- Kite diagrams show both distribution (position) and abundance (width).
- A broad kite indicates high abundance, while narrow or absent sections show low presence or absence.
Sand Dune Zonation - Studland Beach, UK
- Students measured marram grass cover, heather abundance, and slope angle along 100 m transects.
- Data revealed:
- Marram grass abundance peaked in the mid-dunes (stabilizing sand).
- Heather dominated inland grey dunes, where the soil was richer and more stable.
- This study illustrates the interaction between succession and zonation.
- Older dunes further inland supported more complex plant communities.
- Define zonation and give two examples of environmental gradients that cause it.
- What abiotic factors would you measure in a sand dune zonation study?
- Describe how a kite diagram visually represents species distribution and abundance.
- Outline how abiotic gradients influence biotic zonation in a rocky shore ecosystem.


