The Ecological Niche
Ecological niche
An ecological niche is the unique set of abiotic and biotic conditions and resources that a species requires to survive, grow, and reproduce.
- Every species in an ecosystem has a unique ecological niche, its role and position within that system.
- The niche describes the set of abiotic and biotic conditions that determine where and how a species lives, survives, and reproduces.
- It includes all environmental requirements and interactions necessary for an organism’s growth, survival, and reproduction, such as food sources, temperature tolerance, and behavioral adaptations.
Components of a Niche
- Habitat: The physical environment where the species lives (e.g., forest floor, coral reef, tundra).
- Feeding habits: Type of food consumed and method of obtaining it (e.g., carnivore, herbivore, detritivore).
- Temporal activity: When the species is active (diurnal, nocturnal, seasonal).
- Interactions: Symbiotic relationships, competition, and mutualistic partnerships.
- Physiological range: Tolerance limits for temperature, salinity, pH, and humidity.
- Reproductive strategy: Mating behavior, breeding season, and parental investment.
A habitat is the species’ address, while its niche is its occupation, what it does and how it fits into the ecosystem.
When explaining a niche in exams, always include both biotic (feeding, competition, predation) and abiotic (temperature, humidity, soil, light) components.
The Ecological Niche of the Bengal Tiger (Panthera tigris)
- Habitat: Tropical and subtropical forests, mangroves, and grasslands in India, Bangladesh, Nepal, and Bhutan.
- Diet (Food Source): Carnivorous; preys on deer, wild boar, and smaller mammals.
- Hunting Strategy: Ambush predator; hunts alone at night.
- Competitors: Other large carnivores like leopards and dholes (wild dogs).
- Abiotic Factors: Requires dense vegetation for cover, water sources for drinking and cooling, and a temperature range between 20–40°C.
- Predator-prey dynamics: Controls herbivore populations, maintaining ecosystem balance.
- Territorial behavior: Competes with other tigers for territory and mates.
Types of Niches
- Fundamental niche: The full range of environmental conditions under which a species could survive and reproduce.
- Realized niche: The portion of the fundamental niche that a species actually occupies, considering competition and other limiting factors.
Barnacle species on rocky shores illustrate this distinction:
- The species Chthamalus can tolerate a wide range of tidal levels (fundamental niche),
- But due to competition from Balanus, it is confined to higher tidal zones (realized niche).
Competitive Exclusion Principle
- Proposed by G.F. Gause (1934), this principle states that no two species can occupy the exact same niche indefinitely in a stable ecosystem.
- If two species compete for identical resources, the superior competitor will eventually exclude the other.
- Some species avoid this by resource partitioning, i.e., dividing resources spatially or temporally.
- Three North American warbler species feed in the same conifer trees but at different heights.
- This resource partitioning reduces direct competition and allows coexistence.
Population Interactions
- Populations interact in various ways within ecosystems, shaping ecological balance, population sizes, and evolutionary adaptations.
- These interactions drive natural selection, influencing species behavior and survival.
- These interactions create complex food webs and energy flows within ecosystems.
- The major interaction types include herbivory, predation, parasitism, mutualism, disease, and competition.
1. Herbivory
Herbivory
Herbivory is an interaction in which an animal (the herbivore) feeds on a plant or algal species (producer) for energy and nutrients.
- Herbivory occurs when an animal feeds directly on plant material.
- It influences both the plant population (by reducing biomass) and the herbivore population (through food availability).
- Effect on Population Dynamics:
- Overgrazing reduces plant availability, limiting herbivore populations.
- Plants develop chemical defenses, reducing herbivore feeding success.
- The hippopotamus spends nights feeding on grasses and days submerged in rivers.
- It's feeding maintains open grasslands and influences the distribution of aquatic vegetation.
- Ecological impact: Shapes plant community composition.
- Behavioural adaptation: Grazing at night avoids heat stress.
- Evolutionary effect: Selection for digestive systems capable of processing fibrous plants.
Plants evolve defensive adaptations such as thorns, toxins, or thick bark, while herbivores evolve resistance or specialized feeding behaviors.
2. Predation (Predators vs. Prey)
Predation
Predation is an interaction in which one organism (the predator) kills and consumes another organism (the prey) for food.
- Predation is an interaction in which one organism (predator) kills and eats another (prey).
- Predator-prey relationships regulate population sizes through negative feedback loops, maintaining ecological balance.
- Effect on Population Dynamics:
- High parasite loads reduce host survival and reproduction.
- Infected populations may develop partial immunity over generations.
- The lion-zebra relationship in the African savanna:
- Ecological consequence: Controls prey density and prevents overgrazing.
- Behavioural consequence: Zebras form herds for protection and lions hunt cooperatively.
- Evolutionary consequence: Co-evolution of speed, camouflage, and defense strategies in both species.
- Predator-prey population cycles often show a time lag.
- Predator numbers peak after prey populations increase.
3. Parasitism (Parasite vs. Host)
Parasitism
Parasitism is a symbiotic relationship where one organism (the parasite) benefits by living on or within another organism (the host) and obtaining nutrients from it, usually without immediately killing the host.
- Parasitism involves one organism (parasite) benefiting at the expense of another (host).
- The parasite depends on the host for nutrients or habitat, often weakening but not immediately killing it.
- Types:
- Ectoparasites live on the surface (e.g., ticks on dogs).
- Endoparasites live inside the host (e.g., tapeworms in humans).
- Effect on Population Dynamics:
- High parasite loads reduce host survival and reproduction.
- Infected populations may develop partial immunity over generations.
- The malarial parasite (Plasmodium spp.) infects human red blood cells, transmitted by the Anopheles mosquito.
- Ecological effect: Reduces human population health and productivity.
- Behavioral effect: Humans adopt preventive actions (nets, repellents).
- Evolutionary effect: Selection for malaria-resistant genes (e.g., sickle-cell trait).
4. Mutualism (Both Species Benefit)
Mutualism
Mutualism is a symbiotic relationship between two different species in which both organisms benefit from the interaction.
- Mutualism is a symbiotic interaction where both species benefit.
- It enhances survival, reproduction, and carrying capacity for both participants.
- Mutualisms can be obligate (necessary for survival) or facultative (beneficial but not essential).
- Bees collect nectar for food while pollinating flowers, enhancing plant reproduction.
- Ecological effect: Increases productivity and biodiversity.
- Behavioural effect: Species develop interdependent behaviors (e.g., flower recognition).
- Evolutionary effect: Coevolution enhances mutual benefits as floral structures evolve to suit pollinators.
- Effect on Population Dynamics:
- More bees = higher plant reproduction rates.
- More flowering plants = higher bee populations due to nectar availability.
- Coral reefs and zooxanthellae algae:
- The coral provides shelter and CO₂ for the algae.
- The algae photosynthesize and provide glucose for the coral.
- Up to 90% of coral energy comes from this symbiosis.
- Disruption of mutualistic relationships (e.g., coral bleaching) can destabilize entire ecosystems.
5. Disease (Pathogens vs. Hosts)
Disease
Disease is a biological interaction in which a pathogen (virus, bacterium, fungus, or protozoan) infects a host organism, causing harm or death.
- Disease is caused by pathogens (bacteria, viruses, fungi, protozoa) that infect a host, reducing its health and fitness.
- Diseases can act as selective pressures, favoring individuals with natural immunity.
- Effect on Population Dynamics:
- Outbreaks reduce host populations, sometimes leading to local extinction.
- Some populations develop resistance, preventing mass die-offs.
- White-nose syndrome in bats is a fungal disease that disturbs hibernation, leading to starvation and massive population declines.
- Ecological effect: Alters community dynamics and food web balance.
- Behavioural effect: Populations adopt avoidance strategies or develop immunity.
- Evolutionary effect: Acts as a selective pressure favoring resistant individuals.
Always describe how the disease influences population dynamics, e.g., cyclic population drops, increased mortality, or selective resistance.
6. Competition (Interspecific & Intraspecific)
Competition
Competition occurs when two or more organisms require the same limited resource, resulting in a negative effect on all participants’ growth, survival, or reproduction.
- Competition occurs when organisms vie for limited resources such as food, space, water, or mates.
- It can be:
- Intraspecific: Among individuals of the same species.
- Interspecific: Between different species.
- Effect on Population Dynamics:
- Dominant individuals breed more, shaping population genetics.
- Limited resources lead to territorial behaviors and population control.
- Grey squirrels introduced to the UK have outcompeted native red squirrels for food and nesting sites, causing red squirrel decline.
- Ecological effect: Alters species distribution and community composition.
- Behavioural effect: Drives migration, territoriality, or specialization.
- Evolutionary effect: Favors traits that reduce competition (e.g., different feeding times or habitats).
- Define an ecological niche and explain why no two species can occupy the same one.
- Give examples of both fundamental and realized niches.
- Describe one example each of herbivory, predation, parasitism, mutualism, disease, and competition, stating ecological and evolutionary effects.
- Explain how co-evolution operates in predator–prey or parasite–host systems.
- Discuss how population interactions influence ecosystem stability.


