Adaptations of Phloem Sieve Tubes and Companion Cells for Translocation of Sap
- The phloem is a vascular tissue responsible for the translocation of sap, which contains sugars (mainly sucrose), amino acids, and other nutrients.
- The movement occurs from sources (e.g., mature leaves) to sinks (e.g., roots, fruits, and growing tissues).
Structural Adaptations of Phloem Sieve Tubes and Companion Cells
1. Sieve Plates in Phloem Sieve Tube Elements
Sieve tubes
Long, tube-like structures in phloem made up of sieve tube elements connected by sieve plates.
Function
- Sieve plates are specialized end walls between the sieve tube elements.
- These plates have pores that allow the movement of sap from one sieve tube element to the next.

Adaptation
- Pores in sieve plates allow for the unimpeded flow of sap through the phloem.
- This structural adaptation minimizes resistance, making the translocation process more efficient.
2. Reduced Cytoplasm and Organelles in Sieve Tube Elements
Function
- Sieve tube elements have reduced cytoplasm and few organelles to maximize space for sap flow.
- This allows for a continuous column of sap, reducing internal resistance.
Adaptation
- The absence of a nucleus and the reduced number of organelles in sieve tube elements ensures that the cells do not consume the sugars that are being transported.
- This creates an efficient pathway for sap to flow without being interrupted by cellular metabolic processes.
3. Absence of a Nucleus in Sieve Tube Elements
Function

Sieve tube elements lack a nucleus, which would otherwise take up space and potentially compete for energy required for sap movement.
Adaptation
- By not having a nucleus, the sieve tube elements are more efficient at transporting sap because they do not need to allocate resources to maintaining a nucleus.
- Instead, the companion cells provide metabolic support.
Companion cell
Specialized cells in phloem that assist sieve tube elements by providing metabolic support, including energy production.
- Students often forget that sieve tube elements, despite their lack of organelles, are living cells.
- Their survival depends on companion cells.


