Speciation by Splitting of Pre-existing Species
Speciation
Speciation is the evolutionary process through which one species splits into two or more distinct species.
- Speciation is a primary driver of biodiversity, increasing the variety of life.
- It requires reproductive isolation, which allows populations to evolve separately.
- Speciation is distinct from gradual changes within a single species.
- It specifically involves the formation of new species.
Mechanisms of Speciation: The Role of Reproductive Isolation
1. Geographical Isolation (Allopatric Speciation)
- Allopatric speciation occurs when populations are physically separated by barriers such as mountains, rivers, or oceans.
- These barriers prevent interbreeding, allowing populations to evolve independently.
Example: Lava Lizards of the Galápagos Islands
- Lava lizards on the Galápagos Islands are a classic example of allopatric speciation.
- A single species initially colonized the islands, but geographical isolation between the islands prevented interbreeding.
- Over time, genetic drift and natural selection caused the populations on each island to diverge, resulting in multiple unique species.
2. Behavioral or Temporal Isolation (Sympatric Speciation)
- In sympatric speciation, populations live in the same geographical area but become reproductively isolated due to:
- Behavioral Isolation: Differences in mating preferences or courtship behaviors.
In Lake Massoko, two populations of cichlid fish prefer different habitats (shallow vs. deep water), choosing mates based on specific traits.
- Temporal Isolation: Differences in breeding seasons or times.
Winter pine processionary moth populations in Portugal breed during different months, preventing interbreeding.
NoteSympatric speciation is less common than allopatric speciation but demonstrates how species can diverge without physical barriers.
The Process of Speciation: A Step-by-Step Breakdown
1. Separation of Populations:
- Populations become isolated, either:
- Geographically (e.g., rivers, mountains, or islands).
- Reproductively (e.g., differences in behavior or timing).
2. Independent Evolution
- Isolated populations experience different selection pressures, such as:
- Climate changes.
- New predators or competitors.
- Changes in available resources.
Natural selection drives these populations to adapt to their unique environments, causing genetic divergence.
3. Accumulation of Differences:
- Over generations, differences accumulate in:
- Morphology (e.g., body shape or size).
- Behavior (e.g., mating rituals or feeding habits).
- Physiology (e.g., tolerance to environmental factors).
These differences may reduce or eliminate the ability to interbreed.
4. Formation of New Species
- When populations become so distinct that they can no longer interbreed, even if brought back into contact, speciation is complete.
Natural selection acts on genetic variation within each population, gradually shaping traits that are better suited to their specific environment.
Speciation vs. Gradual Evolutionary Change
- While evolution refers to changes in heritable traits over time within a population, speciation specifically involves the splitting of one species into two.
- In contrast, speciation results in distinct species that cannot interbreed.
Antibiotic Resistance: The evolution of antibiotic resistance in bacteria involves gradual changes within a single population and is not an example of speciation.
Common Mistake- Don't confuse gradual changes (e.g., changes in size or coloration) with speciation.
- Speciation requires reproductive isolation and the divergence of populations into distinct species.
Why Speciation Matters
- Speciation plays a central role in increasing biodiversity, contributing to the complexity and stability of ecosystems, and evolutionary innovation.
- Think of speciation as the growth of branches on a tree.
- Each new branch represents a new species, adding to the tree’s complexity and diversity.
- Extinction, in contrast, is like cutting off a branch, reducing the tree’s richness.
Reflection and Broader Implications
- Speciation is a process that continues to shape the diversity of life on Earth. However, it also prompts critical questions:
- How are human activities, such as habitat destruction and climate change, affecting speciation and extinction rates?
- Should humans intervene to prevent species extinction, or is it a natural part of evolution?
Theory of Knowledge Connection:Speciation raises fascinating philosophical questions: How do we define what constitutes a species? Are our definitions influenced by cultural, historical, or scientific perspectives? How does understanding speciation shape our approach to biodiversity and conservation?
Self review- What is the key difference between allopatric and sympatric speciation?
- Why is gradual evolutionary change within a species not considered speciation?
- How does reproductive isolation contribute to the formation of new species?


